Cladoceramus undulatoplicatus (Snowshoe clam)

A fragment of a fossil clam shell in the Fan Bay Deep Shelter tunnels at the National Trust’s White Cliffs of Dover site.

The following photo shows fragments of the clam Cladoceramus undulatoplicatus in the roof of the tunnel above the sheet flint in Photo 2 of “How were flints in chalk formed?”

Fragments of the fossil Cladoceramus undulatoplicatus (Snowshoe Clam).
Image reproduced by kind permission of the National Trust: https//www.nationaltrust.org.uk

It is rare to find such a well-preserved example of a fragment of this clam shell, which is also referred to as a Snowshoe Clam (Everhart, 2017, p. 44). 

The name Snowshoe Clam is appropriate. These clams were large, up to 1 metre across, and they lived on the seabed which was a soft chalky ooze. To prevent sinking into the ooze, they had thin shells compared to many clams, which minimised their weight.  But they also evolved features used in old-fashioned snowshoes which are used to aid walking on soft snow without sinking into it.  The snowshoe and the clam both have an undulating underside, folded like a fan (plicated).  The undulations increase the surface area applied to the soft chalk ooze.  This reduces pressure from the applied weight and the amount of settlement into the soft material. 

The undulations are also reminiscent corrugated metal sheet (“wriggly tin”) which is formed to provide rigidity to an otherwise flimsy sheet material – also applicable to a large thin-shelled clam. 

Normally these fossils are seen in section on vertical exposures, where they show as unremarkable thin wavy lines.  If they are seen in plan, it is usually on a bedding plane exposed on a fallen chalk block at the base of a sea cliff.  These exposures are short lived as they are rapidly broken up by ensuing high tides.  This example is protected from the elements and has been exposed in its current state of preservation since 1941 when the tunnels were excavated.

This fossil species is found in chalk spanning a relatively short period of about 86.3 to 85.7 million years (but also “probably no more than a couple hundred thousand years” has been suggested, (Everhart, 2017)).   The thin layer of rocks that this represents makes it a useful index fossil.  It is used internationally to define the Coniacian/Santonian Stage boundary in the Chalk at 86.3 +/- 0.5 million years.

Andrew Coleman

Rev. 28/04/2026

Reference:

Everhart, M. J. (2017). Oceans of Kansas: A Natural History of the Western Interior Sea (2nd ed.). Indiana University Press.